Ancient Rythulians

The ancient Rythulians inhabited Rythulia before the time of the Wayfarers.

They lacked the ability to fly under their own power.

It's unknown if the ancient Rythulians had "beaks" on their masks like the Ancestors do.

In the Beta
Aside from the six Ancestors, visions suggested that ancient Rythulians appeared pretty much the same as the the Wayfarers do today.

Speculation
The Wayfarers may be reincarnations of ancient Rythulians, now given a chance to atone for their misdeeds.